/*
Theme Name: BiologyInform Science
Theme URI: https://biologyinform.com/
Author: BiologyInform
Description: An editorial bilingual-ready science and biology theme with an expansive reading column, left sidebar, 3D biology hero art, and responsive articles.
Version: 1.4.0
Requires at least: 6.0
Tested up to: 6.8
Requires PHP: 7.4
Text Domain: biologyinform-science
License: GNU General Public License v2 or later
License URI: https://www.gnu.org/licenses/gpl-2.0.html
Tags: blog, education, left-sidebar, custom-logo, custom-menu, featured-images, translation-ready
*/
:root{--ink:#17333b;--muted:#63777a;--teal:#116a70;--deep:#073e49;--green:#6bb58c;--paper:#f6f0e7;--card:#fff;--border:#e1ebe8;--amber:#e7a75e;--shadow:0 10px 40px rgba(13,47,50,.055);--radius:18px}
*,*::before,*::after{box-sizing:border-box}html{scroll-behavior:smooth}body{margin:0;background:var(--paper);color:var(--ink);font:16px/1.66 system-ui,-apple-system,BlinkMacSystemFont,'Segoe UI',sans-serif;-webkit-font-smoothing:antialiased}img{max-width:100%;height:auto}a{color:var(--teal);text-decoration:none}a:hover{text-decoration:underline}button,input,textarea,select{font:inherit}button{cursor:pointer}button:focus-visible,a:focus-visible,input:focus-visible,textarea:focus-visible{outline:3px solid #e8a15b;outline-offset:3px} .screen-reader-text{position:absolute!important;width:1px;height:1px;padding:0;margin:-1px;overflow:hidden;clip:rect(0,0,0,0);white-space:nowrap;border:0}.screen-reader-text:focus{clip:auto!important;width:auto;height:auto;z-index:999;background:white;color:#123;top:6px;left:6px;padding:10px}.container{width:min(100% - 64px,1460px);margin:0 auto}
.topbar{background:#fff;border-bottom:1px solid var(--border)}.topbar-inner{display:flex;align-items:center;gap:20px;min-height:98px}.brand{display:flex;align-items:center;gap:13px;color:var(--deep);min-width:0}.brand:hover{text-decoration:none}.brand-symbol{display:grid;place-items:center;flex:0 0 51px;height:51px;border-radius:16px;background:linear-gradient(135deg,#e2f8ee,#daf3f5);color:#146c67}.brand-mark{width:30px;height:30px}.brand-title{display:block;font-size:25px;font-weight:800;letter-spacing:-.055em;line-height:1.12}.brand-tagline{font-size:12px;color:#698087;line-height:1.4;margin-top:4px}.header-tools{display:flex;gap:15px;align-items:center;margin-left:auto}.header-tools .small-link{font-size:13px;font-weight:700;color:#456b70;white-space:nowrap}.header-search{display:flex;align-items:center;gap:8px;max-width:232px;background:#f3f7f7;border:1px solid var(--border);border-radius:999px;padding:5px 11px}.header-search input{border:0;outline:none;background:transparent;width:100%;font-size:13px;color:var(--ink)}.header-search button{border:0;background:none;padding:0;color:var(--teal);display:grid;place-items:center}.header-search svg{width:18px;height:18px}.mobile-menu-button{display:none;border:1px solid var(--border);border-radius:12px;background:white;color:var(--deep);padding:8px 11px;font-weight:700}
.site-shell{display:grid;grid-template-columns:272px minmax(0,1fr);gap:30px;align-items:start;padding-top:30px;padding-bottom:64px}.sidebar{position:sticky;top:22px;align-self:start}.sidebar-inner{background:white;border:1px solid var(--border);border-radius:var(--radius);padding:22px 17px;box-shadow:var(--shadow)}.sidebar-kicker{margin:3px 13px 15px;font-size:10px;letter-spacing:.18em;text-transform:uppercase;color:#829797;font-weight:800}.sidebar .widget{margin-bottom:29px}.sidebar .widget:last-child{margin-bottom:0}.sidebar .widget-title{font-size:11px;letter-spacing:.14em;text-transform:uppercase;color:#829797;margin:0 13px 10px}.sidebar ul,.sidebar .menu{padding:0;margin:0;list-style:none}.sidebar li{list-style:none;margin:0 0 3px}.sidebar li a{display:block;color:#36575d;padding:9px 12px;font-size:13.5px;font-weight:600;line-height:1.32;border-radius:10px;transition:background .16s,color .16s}.sidebar li a:hover,.sidebar li.current-menu-item>a,.sidebar li.current-cat>a{background:#e8f5f0;color:#096167;text-decoration:none}.sidebar li ul{margin-left:12px;border-left:1px solid #e2eae7;padding-left:7px}.sidebar li li a{font-size:12px;padding:7px 10px}.sidebar-nav-label{padding:0 13px 8px;font-size:11px;letter-spacing:.15em;text-transform:uppercase;color:#859997;font-weight:800}.sidebar-bottom{border-radius:13px;margin-top:19px;background:linear-gradient(140deg,#eaf5f0,#e8f1f8);padding:16px}.sidebar-bottom strong{display:block;font-size:13px;color:#155a5b}.sidebar-bottom p{font-size:12px;color:#55757a;margin:5px 0 0}.sidebar-bottom a{font-size:12px;font-weight:800}
.primary-content{min-width:0}.hero{position:relative;min-height:380px;overflow:hidden;border-radius:24px;background:#083a43 url('assets/images/biology-hero.webp') center center/cover no-repeat;isolation:isolate;display:flex;align-items:center;box-shadow:var(--shadow)}.hero::before{content:"";position:absolute;inset:0;z-index:-1;background:linear-gradient(90deg,rgba(2,44,54,.96) 0%,rgba(2,44,54,.78) 29%,rgba(3,48,53,.15) 73%,rgba(2,29,37,.05) 100%)}.hero-body{padding:48px clamp(25px,5vw,66px);max-width:670px}.eyebrow{font-size:11px;text-transform:uppercase;letter-spacing:.18em;color:#b8e8cf;font-weight:850}.hero h1{color:#fff;font-size:clamp(36px,4.4vw,64px);letter-spacing:-.055em;line-height:1.055;margin:13px 0 17px;font-weight:800}.hero p{color:#e1f2ee;font-size:clamp(14px,1.45vw,17px);max-width:480px;line-height:1.68;margin:0 0 27px}.hero-actions{display:flex;gap:11px;flex-wrap:wrap}.btn{display:inline-flex;gap:8px;align-items:center;justify-content:center;padding:12px 19px;border-radius:999px;font-weight:800;font-size:13px;transition:transform .15s}.btn:hover{text-decoration:none;transform:translateY(-2px)}.btn-primary{color:#0a3e44;background:#cdf1d9}.btn-ghost{border:1px solid #a9c8c7;color:white;background:rgba(255,255,255,.06)}.btn-ghost:hover{color:white}
.mini-intro{margin:26px 0 30px;padding:18px 23px;background:white;border:1px solid var(--border);border-radius:15px;display:flex;gap:16px;align-items:center;box-shadow:var(--shadow)}.mini-intro-icon{font-size:23px;display:grid;place-items:center;flex:0 0 42px;width:42px;height:42px;background:#e7f5ef;border-radius:12px}.mini-intro p{margin:0;color:#536d70;font-size:14px}.mini-intro strong{color:#1e5556}.section-header{display:flex;justify-content:space-between;align-items:end;gap:20px;margin:38px 0 17px}.section-header h2{font-size:clamp(23px,2.3vw,31px);letter-spacing:-.04em;line-height:1.15;margin:0}.section-header p{font-size:13px;color:#738788;margin:8px 0 0}.section-action{font-size:13px;font-weight:750;white-space:nowrap}.topic-grid{display:grid;grid-template-columns:repeat(4,minmax(0,1fr));gap:13px}.topic-card{padding:20px 17px;display:flex;min-height:160px;flex-direction:column;align-items:start;justify-content:space-between;background:#fff;border-radius:17px;border:1px solid var(--border);position:relative;overflow:hidden;box-shadow:var(--shadow);color:var(--deep)}.topic-card:after{content:"";position:absolute;right:-25px;bottom:-45px;width:120px;height:120px;background:var(--topic-accent,#dcf5e9);border-radius:50%;opacity:.65}.topic-card:hover{text-decoration:none;transform:translateY(-3px);transition:transform .2s}.topic-emoji{position:relative;z-index:1;font-size:30px;line-height:1}.topic-title{position:relative;z-index:1;font-weight:800;font-size:15px;line-height:1.25}.topic-more{position:relative;z-index:1;display:block;font-size:11px;color:#547d77}.story-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:17px}.story-card{background:#fff;border:1px solid var(--border);border-radius:18px;overflow:hidden;box-shadow:var(--shadow);display:flex;flex-direction:column;height:100%}.story-card-cover{display:block;position:relative;aspect-ratio:1.6;background:linear-gradient(135deg,#e0ede5,#d5e8f2);overflow:hidden}.story-card-cover img{width:100%;height:100%;object-fit:cover;display:block;transition:transform .25s}.story-card:hover img{transform:scale(1.035)}.story-cover-placeholder{position:absolute;inset:0;display:grid;place-items:center;font-size:52px;color:#2b6771;background:radial-gradient(circle at 70% 25%,#f8fff9,transparent 40%),linear-gradient(125deg,#d9f1e7,#d4e7f1)}.story-card-body{padding:17px 18px 19px;display:flex;flex-direction:column;flex:1}.story-meta{font-size:10.5px;color:#6a8c87;text-transform:uppercase;font-weight:850;letter-spacing:.09em}.story-card h3{line-height:1.36;font-size:17px;margin:8px 0 8px;letter-spacing:-.024em}.story-card h3 a{color:#193d44}.story-card h3 a:hover{color:#0e767a}.story-card p{margin:0 0 15px;font-size:12.5px;line-height:1.7;color:#728586;flex:1}.read-link{font-size:12px;font-weight:850}.empty-note{background:white;border:1px dashed #c1d7d5;border-radius:17px;padding:25px;color:#6d8685;grid-column:1/-1}.feature-strip{margin-top:34px;border-radius:19px;background:linear-gradient(115deg,#124d55,#0d7476);color:white;padding:25px 30px;display:flex;align-items:center;justify-content:space-between;gap:18px}.feature-strip .overline{color:#bbdfcf;text-transform:uppercase;letter-spacing:.16em;font-size:10px;font-weight:800}.feature-strip h2{color:white;margin:5px 0 5px;font-size:24px;letter-spacing:-.03em}.feature-strip p{color:#e1f1ea;margin:0;font-size:13px}.feature-strip a{white-space:nowrap;background:#ddf5dc;color:#0f4a50;font-weight:800;padding:11px 18px;border-radius:999px;font-size:12px}.feature-strip a:hover{text-decoration:none}
.content-panel{background:#fff;border:1px solid var(--border);border-radius:22px;padding:clamp(23px,4vw,56px);box-shadow:var(--shadow)}.crumbs{display:flex;flex-wrap:wrap;gap:9px;color:#79908e;font-size:12px;margin:0 0 24px}.crumbs span{color:#b1c2be}.entry-title,.archive-title{font-size:clamp(31px,3.6vw,51px);letter-spacing:-.055em;line-height:1.14;margin:4px 0 17px;font-weight:800;color:#123f47;overflow-wrap:anywhere}.entry-byline{display:flex;align-items:center;gap:10px;flex-wrap:wrap;color:#829493;font-size:13px;margin:0 0 23px}.entry-featured{margin:0 0 25px;border-radius:15px;overflow:hidden}.entry-featured img{width:100%;max-height:540px;object-fit:cover;display:block}.entry-content{font-size:18px;line-height:1.82;color:#334e54;overflow-wrap:break-word}.entry-content p{margin:0 0 1.35em}.entry-content h2,.entry-content h3{color:#124a53;letter-spacing:-.035em;line-height:1.25;margin:1.7em 0 .65em}.entry-content h2{font-size:1.65em}.entry-content h3{font-size:1.29em}.entry-content img{border-radius:12px}.entry-content figure{max-width:100%}.entry-content ul,.entry-content ol{padding-left:1.5em}.entry-content blockquote{border-left:4px solid #88c9ad;margin:26px 0;padding:4px 23px;background:#f5faf7;border-radius:0 12px 12px 0;color:#456b66}.entry-content a{text-decoration:underline;text-underline-offset:2px}.entry-content :is(iframe,video){max-width:100%}.entry-content .wp-block-embed__wrapper{position:relative;max-width:100%}.entry-content .wp-block-embed.is-type-video .wp-block-embed__wrapper{aspect-ratio:16/9}.entry-content .wp-block-embed.is-type-video iframe{width:100%;height:100%;position:absolute;inset:0;border:0}.entry-content audio{width:100%;margin:12px 0}.entry-content table{border-collapse:collapse;width:100%;display:block;overflow-x:auto;font-size:15px}.entry-content th,.entry-content td{border:1px solid var(--border);padding:10px 12px}.entry-content th{background:#f1f8f5}.post-tags{display:flex;gap:8px;flex-wrap:wrap;margin-top:30px}.post-tags a{padding:7px 11px;background:#edf6f2;border-radius:99px;font-size:12px;color:#306d65}.page-links{margin:24px 0}.navigation.post-navigation{border-top:1px solid var(--border);margin-top:35px;padding-top:21px}.nav-links{display:flex;justify-content:space-between;gap:12px;font-size:13px}.nav-links a{font-weight:700}.archive-desc{font-size:16px;color:#78908d;line-height:1.7;margin:0 0 25px}.pager{margin:28px 0}.page-numbers{display:inline-block;padding:8px 12px;border-radius:9px;background:white;border:1px solid #d9e9e6;margin:0 4px 7px 0}.page-numbers.current{color:white;background:#176c71;border-color:#176c71}.not-found{text-align:center;padding:70px 30px}.not-found h1{font-size:48px}.site-footer{border-top:1px solid var(--border);background:#fff}.footer-inner{display:flex;justify-content:space-between;gap:15px;padding:29px 0;font-size:12px;color:#748684}.footer-inner strong{color:#23545a}.footer-inner a{color:#41676a}.wp-caption{max-width:100%}.wp-caption-text,.gallery-caption{font-size:13px;color:#81918f}.alignleft{float:left;margin:5px 24px 16px 0}.alignright{float:right;margin:5px 0 16px 24px}.aligncenter{display:block;margin:15px auto}.sticky{border:1px solid #95cab9}.bypostauthor{font-weight:inherit}.comment-list{padding-left:22px}.comment-list li{margin-bottom:18px}.comment-respond input:not([type=submit]),.comment-respond textarea{max-width:100%;width:100%;border:1px solid var(--border);padding:12px;border-radius:10px}.comment-respond input[type=submit]{border:0;color:#fff;background:#12646b;padding:10px 17px;border-radius:9px}
@media(max-width:1180px){.container{width:min(100% - 40px,1460px)}.site-shell{grid-template-columns:244px minmax(0,1fr);gap:20px}.topic-grid{grid-template-columns:repeat(2,minmax(0,1fr))}.story-grid{grid-template-columns:repeat(2,minmax(0,1fr))}}
@media(max-width:800px){.container{width:min(100% - 28px,1460px)}.topbar-inner{min-height:76px}.brand-symbol{height:43px;flex-basis:43px;border-radius:11px}.brand-title{font-size:21px}.brand-tagline{font-size:10px}.header-search{max-width:155px}.mobile-menu-button{display:block}.site-shell{display:flex;flex-direction:column;padding-top:17px;gap:16px}.sidebar{display:none;position:static;width:100%}.sidebar.is-open{display:block}.sidebar-inner{padding:17px}.primary-content{width:100%}.hero{min-height:340px}.hero-body{padding:33px 25px}.hero::before{background:linear-gradient(90deg,rgba(3,41,49,.96),rgba(3,41,49,.66) 62%,rgba(3,41,49,.20))}}
@media(max-width:540px){.header-tools .small-link,.header-search{display:none}.topbar-inner{gap:8px}.hero{min-height:370px;background-position:65% center}.hero h1{font-size:38px}.hero p{font-size:14px}.mini-intro{align-items:flex-start;padding:15px}.section-header{margin-top:30px}.topic-grid,.story-grid{grid-template-columns:1fr 1fr;gap:10px}.topic-card{min-height:136px;padding:15px}.topic-title{font-size:13px}.topic-emoji{font-size:26px}.story-grid{grid-template-columns:1fr}.feature-strip{align-items:flex-start;flex-direction:column;padding:24px}.feature-strip h2{font-size:22px}.content-panel{border-radius:16px;padding:22px 18px}.entry-content{font-size:16.5px}.footer-inner{flex-direction:column}.entry-title{font-size:33px}}
@media(prefers-reduced-motion:reduce){*,*::before,*::after{scroll-behavior:auto!important;transition:none!important}}

.brand-copy{color:var(--deep)}.brand-copy:hover{text-decoration:none}.brand-custom-logo{display:block;max-width:170px}.brand-custom-logo img{max-height:52px;width:auto;display:block}.brand-symbol:hover{text-decoration:none}.content-panel form[role=search] {display:flex;gap:9px;align-items:center;flex-wrap:wrap;margin:14px 0 30px}.content-panel form[role=search] label{width:100%;font-size:13px;color:var(--muted)}.content-panel form[role=search] input{padding:10px 12px;flex:1;min-width:150px;border:1px solid var(--border);border-radius:9px}.content-panel form[role=search] button{background:var(--teal);border:0;color:white;padding:10px 18px;border-radius:9px}.wp-block-video video{width:100%}.wp-block-audio audio{width:100%}


/* === BiologyInform Science 1.1 pastel refresh === */
:root{--paper-blue:#edf5f8;--sidebar-bg-1:#dce9e9;--sidebar-bg-2:#eef4f1;--sidebar-border:#c7d8d7;--sidebar-accent:#8fb7b3}
body{background:linear-gradient(180deg,#f8f2e7 0%,#eef5f8 52%,#f6f2ea 100%);position:relative;overflow-x:hidden}
body::before{content:"";position:fixed;inset:0;pointer-events:none;opacity:.34;background:
 radial-gradient(170px 170px at 8% 12%, rgba(147,188,201,.16) 0 2%, transparent 2.8%, transparent 10%, rgba(147,188,201,.10) 10.5%, transparent 11.2%),
 radial-gradient(220px 220px at 87% 22%, rgba(132,170,184,.11) 0 2%, transparent 2.8%, transparent 10%, rgba(132,170,184,.08) 10.5%, transparent 11.1%),
 radial-gradient(250px 250px at 22% 78%, rgba(121,176,153,.10) 0 1.8%, transparent 2.5%, transparent 10%, rgba(121,176,153,.08) 10.5%, transparent 11%),
 radial-gradient(600px 220px at 15% 35%, rgba(255,255,255,.28), transparent 62%),
 radial-gradient(520px 220px at 88% 75%, rgba(183,211,220,.18), transparent 62%),
 linear-gradient(180deg, rgba(255,255,255,.10), rgba(255,255,255,.03));
filter:blur(.2px)}
.topbar{background:rgba(255,251,246,.84);backdrop-filter:blur(8px);border-bottom:1px solid rgba(199,216,215,.85)}
.header-search{background:rgba(255,255,255,.75)}
.sidebar-inner{background:linear-gradient(180deg,var(--sidebar-bg-1) 0%, var(--sidebar-bg-2) 100%);border:1px solid var(--sidebar-border);box-shadow:0 18px 48px rgba(37,70,73,.10), inset 0 1px 0 rgba(255,255,255,.7);position:relative;overflow:hidden}
.sidebar-inner::before{content:"";position:absolute;inset:-30% -12% auto auto;width:190px;height:190px;border-radius:50%;background:radial-gradient(circle at 40% 40%, rgba(255,255,255,.42), rgba(255,255,255,0) 68%);pointer-events:none}
.sidebar-inner::after{content:"";position:absolute;left:-22px;bottom:24px;width:130px;height:130px;border-radius:50%;border:1px solid rgba(112,154,160,.16);box-shadow:40px -42px 0 0 rgba(255,255,255,0), 16px 14px 0 0 rgba(112,154,160,.10), 56px 36px 0 0 rgba(112,154,160,.08);opacity:.8;pointer-events:none}
.sidebar-nav-label,.sidebar-kicker,.sidebar .widget-title{color:#56757a}
.sidebar li a{color:#29474f}
.sidebar li a:hover,.sidebar li.current-menu-item>a,.sidebar li.current-cat>a{background:rgba(255,255,255,.68);color:#173f47;box-shadow:inset 0 0 0 1px rgba(160,190,194,.34)}
.sidebar-bottom{background:rgba(255,255,255,.48);border:1px solid rgba(168,191,193,.46);border-radius:14px;padding:14px 14px 13px}
.primary-content>.content-panel,.story-card,.topic-card,.mini-intro,.content-panel{background:rgba(255,255,255,.88);backdrop-filter:blur(2px)}
.hero{border:1px solid rgba(214,226,225,.86)}
@media (max-width: 860px){body::before{opacity:.24}.sidebar-inner{background:linear-gradient(180deg,#e3eeed 0%,#f2f6f3 100%)}}


/* 1.3: hero image is an HTML element to avoid CSS background optimization and WebP compatibility issues. */
.hero {isolation:isolate;background-color:#164d52}
.hero-art {position:absolute;inset:0;width:100%;height:100%;display:block;object-fit:cover;object-position:center center;z-index:0;pointer-events:none}
.hero::before {z-index:1;pointer-events:none;background:linear-gradient(90deg,rgba(2,44,54,.94) 0%,rgba(2,44,54,.74) 30%,rgba(3,48,53,.13) 72%,rgba(2,29,37,.03) 100%)}
.hero-body {position:relative;z-index:2}
@media(max-width:800px){.hero::before {background:linear-gradient(90deg,rgba(3,41,49,.94),rgba(3,41,49,.68) 62%,rgba(3,41,49,.16))}}


/* 1.4: soft sea-glass left navigation — distinct panel with dark accessible text. */
.sidebar-inner {
  isolation:isolate;
  background:linear-gradient(160deg,#afd1d4 0%,#c8e0de 46%,#d3e9e4 100%);
  border:1px solid #9ebfc2;
  color:#183d48;
  box-shadow:0 16px 36px rgba(39,85,92,.13),inset 0 1px 0 rgba(255,255,255,.55);
}
.sidebar-inner::before {
  content:"";
  position:absolute;
  inset:0;
  width:auto;
  height:auto;
  border:0;
  border-radius:inherit;
  opacity:.26;
  background-image:radial-gradient(circle at 83% 14%,transparent 0 31px,rgba(56,126,138,.15) 32px,transparent 34px),radial-gradient(circle at 35% 72%,transparent 0 40px,rgba(50,113,129,.12) 41px,transparent 43px),radial-gradient(circle at 28% 75%,rgba(255,255,255,.23) 0 9px,transparent 11px);
  pointer-events:none;
  z-index:0;
}
.sidebar-inner::after{display:none}
.sidebar-inner > *{position:relative;z-index:1}
.sidebar-nav-label,.sidebar-kicker,.sidebar .widget-title{color:#3b646c;text-shadow:none}
.sidebar li a{color:#1b4650;font-weight:650}
.sidebar li a:hover,.sidebar li.current-menu-item > a,.sidebar li.current-cat > a,
.sidebar li.current-menu-ancestor > a,.sidebar li.current_page_item > a {
  background:#e6f7ed;
  color:#124650;
  box-shadow:inset 0 0 0 1px rgba(126,170,154,.45),0 2px 8px rgba(36,84,87,.07);
  text-decoration:none;
}
.sidebar li ul{border-left-color:#9cbec1}
.sidebar .widget,.sidebar .widget p,.sidebar .widget li,.sidebar .widget .textwidget{color:#234e55}
.sidebar .widget a{color:#17464d}
.sidebar .widget a:hover{color:#0d565d}
.sidebar-bottom{background:rgba(247,254,251,.53);border:1px solid rgba(122,166,168,.44);color:#204e52}
.sidebar-bottom strong{color:#135b5e}
.sidebar-bottom p{color:#456e73}
.sidebar-bottom a{color:#0e6568}
.feature-strip {background:linear-gradient(115deg,#124d55,#0d7476);color:white}
@media(max-width:860px){.sidebar-inner{background:linear-gradient(160deg,#afd1d4,#d3e9e4)}}
/* Override earlier background shorthand safely: the hero artwork is a real image element. */
.hero {background:#164d52;position:relative}
.hero-art{display:block;position:absolute;inset:0;width:100%;height:100%;object-fit:cover;object-position:65% center;z-index:0}
.hero-body{position:relative;z-index:2}
.hero::before{z-index:1;pointer-events:none}